Data files contain the simulated mean daily environmental conditions experienced by particles (which can be assumed to be planktonic larval alewife or yellow perch) in Lake Michigan during March 15 - July 31 of past and future years. Simulations were generated by integrating a series of climatic, hydrodynamic, biogeochemical, and Lagrangian particle dispersion (LPD) models. For each year, 42,765 particles (i.e., each representing a theoretical larval fish) were released from coastal and nearshore locations and tracked sub-hourly for 50-days following their release. Each year's dataset summarizes individual particle experiences (e.g., light, zooplankton prey, temperature) into daily averages.