The northward migration of the Mendocino triple junction (MTJ) drives a fundamental plate boundary transformation from convergence to translation; producing a series of strike-slip faults, that become the San Andreas plate boundary. We find that the 3-D structure of the Pacific plate lithosphere in the vicinity of the MTJ controls the location of San Andreas plate boundary formation. At the time of initiation of the Pacific-North America plate boundary (∼30 Ma), the sequential interaction with the western margin of North America of the Pioneer Fracture Zone, soon followed by the Mendocino Fracture Zone, led to the capture of a small segment of partially subducted Farallon lithosphere by the Pacific plate, termed the Pioneer Fragment (PF). Since that time, the PF has translated with the Pacific Plate along the western margin of North America. Recently developed, high-resolution seismic-tomographic imagery of northern California indicates that (a) the PF is extant, occupying the western half of the slab window, immediately south of the MTJ; (b) the eastern edge of the PF lies beneath the newly forming Maacama fault system, which develops to become the locus for the primary plate boundary structure after approximately 6–10 Ma; and (c) the location of the translating PF adjacent to the asthenosphere of the slab window generates a shear zone within and below the crust that develops into the plate boundary faults. As a result, the San Andreas plate boundary forms interior to the western margin of North America, rather than at its western edge.
