Mark Haroldson
Mark Haroldson is a USGS Supervisory Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT.
Research Interest
Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Education and Certifications
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Science and Products
Filter Total Items: 81
Influence of whitebark pine decline on fall habitat use and movements of grizzly bears in the Greater Yellowstone Ecosystem
When abundant, seeds of the high-elevation whitebark pine (WBP; Pinus albicaulis) are an important fall food for grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem. Rates of bear mortality and bear/human conflicts have been inversely associated with WBP productivity. Recently, mountain pine beetles (Dendroctonus ponderosae) have killed many cone-producing WBP trees. We used fall (15
Authors
Cecily M. Costello, Frank T. van Manen, Mark A. Haroldson, Michael R. Ebinger, Steven L. Cain, Kerry A. Gunther, Daniel D. Bjornlie
Re-evaluation of Yellowstone grizzly bear population dynamics not supported by empirical data: response to Doak & Cutler
Doak and Cutler critiqued methods used by the Interagency Grizzly Bear Study Team (IGBST) to estimate grizzly bear population size and trend in the Greater Yellowstone Ecosystem. Here, we focus on the premise, implementation, and interpretation of simulations they used to support their arguments. They argued that population increases documented by IGBST based on females with cubs-of-the-year were
Authors
Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Richard B. Harris, Megan D. Higgs, Steve Cherry, Gary C. White, Charles C. Schwartz
Methods to estimate distribution and range extent of grizzly bears in the Greater Yellowstone Ecosystem
The distribution of the Greater Yellowstone Ecosystem grizzly bear (Ursus arctos) population has expanded into areas unoccupied since the early 20th century. Up-to-date information on the area and extent of this distribution is crucial for federal, state, and tribal wildlife and land managers to make informed decisions regarding grizzly bear management. The most recent estimate of grizzly bear dis
Authors
Mark A. Haroldson, Charles C. Schwartz, Daniel J. Thompson, Daniel D. Bjornlie, Kerry A. Gunther, Steven L. Cain, Daniel B. Tyers, Kevin L. Frey, Bryan C. Aber
Contrasting past and current numbers of bears visiting Yellowstone cutthroat trout streams
Spawning cutthroat trout (Oncorhynchus clarkii bouvieri) were historically abundant within tributary streams of Yellowstone Lake within Yellowstone National Park and were a highly digestible source of energy and protein for Yellowstone’s grizzly bears (Ursus arctos) and black bears (U. americanus). The cutthroat trout population has subsequently declined since the introduction of non-native lake t
Authors
Mark A. Haroldson, Charles C. Schwartz, Justin E. Teisberg, Kerry A. Gunther, Jennifer K. Fortin, Charles T. Robbins
Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2013. The report also contains a summary of nuisance grizzly bear management actions.
Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem
Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE) are opportunistic omnivores that eat a great diversity of plant and animal species. Changes in climate may affect regional vegetation, hydrology, insects, and fire regimes, likely influencing the abundance, range, and elevational distribution of the plants and animals consumed by GYE grizzly bears. Determining the dietary brea
Authors
Kerry A. Gunther, Rebecca Shoemaker, Kevin L. Frey, Mark A. Haroldson, Steven L. Cain, Frank T. van Manen, Jennifer K. Fortin
Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee
The Yellowstone grizzly bear (Ursus arctos) was listed as a threatened species in 1975 (Federal Register 40 FR:31734-31736). Since listing, recovery efforts have focused on increasing population size, improving habitat security, managing bear mortalities, and reducing bear-human conflicts. The Interagency Grizzly Bear Committee (IGBC; partnership of federal and state agencies responsible for grizz
Authors
Frank T. van Manen, Cecily M. Costello, Mark A. Haroldson, Daniel D. Bjornlie, Michael R. Ebinger, Kerry A. Gunther, Mary Frances Mahalovich, Daniel J. Thompson, Megan D. Higgs, Kathryn M. Irvine, Kristin Legg, Daniel B. Tyers, Lisa Landenburger, Steven L. Cain, Kevin L. Frey, Bryan C. Aber, Charles C. Schwartz
Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year
Mark-resight designs for estimation of population abundance are common and attractive to researchers. However, inference from such designs is very limited when faced with sparse data, either from a low number of marked animals, a low probability of detection, or both. In the Greater Yellowstone Ecosystem, yearly mark-resight data are collected for female grizzly bears with cubs-of-the-year (FCOY),
Authors
Megan D. Higgs, William A. Link, Gary C. White, Mark A. Haroldson, Daniel D. Bjornlie
Omnivory and the terrestrial food web: Yellowstone grizzly beard diets
No abstract available.
Authors
Charles C. Schwartz, Mark A. Haroldson, Kerry A. Gunther, Charles T. Robbins
Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem
The Greater Yellowstone Ecosystem (GYE) has experienced changes in the distribution and availability of grizzly bear (Ursus arctos) food resources in recent decades. The decline of ungulates, fish, and whitebark pine seeds (Pinus albicaulis) has prompted questions regarding their ability to adapt. We examined body composition and diet of grizzly bears using bioelectrical impedance and stable isoto
Authors
Charles C. Schwartz, Jennifer K. Fortin, Justin E. Teisberg, Mark A. Haroldson, Christopher Servheen, Charles T. Robbins, Frank T. van Manen
Population fragmentation and inter-ecosystem movements of grizzly bears in Western Canada and the Northern United States
Population fragmentation compromises population viability, reduces a species ability to respond to climate change, and ultimately may reduce biodiversity. We studied the current state and potential causes of fragmentation in grizzly bears over approximately 1,000,000 km 2 of western Canada, the northern United States (US), and southeast Alaska. We compiled much of our data from projects undertake
Authors
M.F. Proctor, David Paetkau, B. N. McLellan, G.B. Stenhouse, K.C. Kendall, R.D. Mace, W.F. Kasworm, C. Servheen, C.L. Lausen, M.L. Gibeau, W.L. Wakkinen, M.A. Haroldson, G. Mowat, C.D. Apps, L.M. Ciarniello, R.M.R. Barclay, M.S. Boyce, C.C. Schwartz, C. Strobeck
Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana
We estimated grizzly bear (Ursus arctos) population vital rates and trend for the Northern Continental Divide Ecosystem (NCDE), Montana, between 2004 and 2009 by following radio-collared females and observing their fate and reproductive performance. Our estimates of dependent cub and yearling survival were 0.612 (95% CI = 0.300–0.818) and 0.682 (95% CI = 0.258–0.898). Our estimates of subadult and
Authors
R.D. Mace, D.W. Carney, T. Chilton-Radandt, S.A. Courville, M.A. Haroldson, R.B. Harris, J. Jonkel, B. McLellan, M. Madel, T.L. Manley, C.C. Schwartz, C. Servheen, G. Stenhouse, J.S. Waller, E. Wenum
Science and Products
Filter Total Items: 81
Influence of whitebark pine decline on fall habitat use and movements of grizzly bears in the Greater Yellowstone Ecosystem
When abundant, seeds of the high-elevation whitebark pine (WBP; Pinus albicaulis) are an important fall food for grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem. Rates of bear mortality and bear/human conflicts have been inversely associated with WBP productivity. Recently, mountain pine beetles (Dendroctonus ponderosae) have killed many cone-producing WBP trees. We used fall (15
Authors
Cecily M. Costello, Frank T. van Manen, Mark A. Haroldson, Michael R. Ebinger, Steven L. Cain, Kerry A. Gunther, Daniel D. Bjornlie
Re-evaluation of Yellowstone grizzly bear population dynamics not supported by empirical data: response to Doak & Cutler
Doak and Cutler critiqued methods used by the Interagency Grizzly Bear Study Team (IGBST) to estimate grizzly bear population size and trend in the Greater Yellowstone Ecosystem. Here, we focus on the premise, implementation, and interpretation of simulations they used to support their arguments. They argued that population increases documented by IGBST based on females with cubs-of-the-year were
Authors
Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Richard B. Harris, Megan D. Higgs, Steve Cherry, Gary C. White, Charles C. Schwartz
Methods to estimate distribution and range extent of grizzly bears in the Greater Yellowstone Ecosystem
The distribution of the Greater Yellowstone Ecosystem grizzly bear (Ursus arctos) population has expanded into areas unoccupied since the early 20th century. Up-to-date information on the area and extent of this distribution is crucial for federal, state, and tribal wildlife and land managers to make informed decisions regarding grizzly bear management. The most recent estimate of grizzly bear dis
Authors
Mark A. Haroldson, Charles C. Schwartz, Daniel J. Thompson, Daniel D. Bjornlie, Kerry A. Gunther, Steven L. Cain, Daniel B. Tyers, Kevin L. Frey, Bryan C. Aber
Contrasting past and current numbers of bears visiting Yellowstone cutthroat trout streams
Spawning cutthroat trout (Oncorhynchus clarkii bouvieri) were historically abundant within tributary streams of Yellowstone Lake within Yellowstone National Park and were a highly digestible source of energy and protein for Yellowstone’s grizzly bears (Ursus arctos) and black bears (U. americanus). The cutthroat trout population has subsequently declined since the introduction of non-native lake t
Authors
Mark A. Haroldson, Charles C. Schwartz, Justin E. Teisberg, Kerry A. Gunther, Jennifer K. Fortin, Charles T. Robbins
Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2013. The report also contains a summary of nuisance grizzly bear management actions.
Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem
Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE) are opportunistic omnivores that eat a great diversity of plant and animal species. Changes in climate may affect regional vegetation, hydrology, insects, and fire regimes, likely influencing the abundance, range, and elevational distribution of the plants and animals consumed by GYE grizzly bears. Determining the dietary brea
Authors
Kerry A. Gunther, Rebecca Shoemaker, Kevin L. Frey, Mark A. Haroldson, Steven L. Cain, Frank T. van Manen, Jennifer K. Fortin
Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee
The Yellowstone grizzly bear (Ursus arctos) was listed as a threatened species in 1975 (Federal Register 40 FR:31734-31736). Since listing, recovery efforts have focused on increasing population size, improving habitat security, managing bear mortalities, and reducing bear-human conflicts. The Interagency Grizzly Bear Committee (IGBC; partnership of federal and state agencies responsible for grizz
Authors
Frank T. van Manen, Cecily M. Costello, Mark A. Haroldson, Daniel D. Bjornlie, Michael R. Ebinger, Kerry A. Gunther, Mary Frances Mahalovich, Daniel J. Thompson, Megan D. Higgs, Kathryn M. Irvine, Kristin Legg, Daniel B. Tyers, Lisa Landenburger, Steven L. Cain, Kevin L. Frey, Bryan C. Aber, Charles C. Schwartz
Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year
Mark-resight designs for estimation of population abundance are common and attractive to researchers. However, inference from such designs is very limited when faced with sparse data, either from a low number of marked animals, a low probability of detection, or both. In the Greater Yellowstone Ecosystem, yearly mark-resight data are collected for female grizzly bears with cubs-of-the-year (FCOY),
Authors
Megan D. Higgs, William A. Link, Gary C. White, Mark A. Haroldson, Daniel D. Bjornlie
Omnivory and the terrestrial food web: Yellowstone grizzly beard diets
No abstract available.
Authors
Charles C. Schwartz, Mark A. Haroldson, Kerry A. Gunther, Charles T. Robbins
Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem
The Greater Yellowstone Ecosystem (GYE) has experienced changes in the distribution and availability of grizzly bear (Ursus arctos) food resources in recent decades. The decline of ungulates, fish, and whitebark pine seeds (Pinus albicaulis) has prompted questions regarding their ability to adapt. We examined body composition and diet of grizzly bears using bioelectrical impedance and stable isoto
Authors
Charles C. Schwartz, Jennifer K. Fortin, Justin E. Teisberg, Mark A. Haroldson, Christopher Servheen, Charles T. Robbins, Frank T. van Manen
Population fragmentation and inter-ecosystem movements of grizzly bears in Western Canada and the Northern United States
Population fragmentation compromises population viability, reduces a species ability to respond to climate change, and ultimately may reduce biodiversity. We studied the current state and potential causes of fragmentation in grizzly bears over approximately 1,000,000 km 2 of western Canada, the northern United States (US), and southeast Alaska. We compiled much of our data from projects undertake
Authors
M.F. Proctor, David Paetkau, B. N. McLellan, G.B. Stenhouse, K.C. Kendall, R.D. Mace, W.F. Kasworm, C. Servheen, C.L. Lausen, M.L. Gibeau, W.L. Wakkinen, M.A. Haroldson, G. Mowat, C.D. Apps, L.M. Ciarniello, R.M.R. Barclay, M.S. Boyce, C.C. Schwartz, C. Strobeck
Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana
We estimated grizzly bear (Ursus arctos) population vital rates and trend for the Northern Continental Divide Ecosystem (NCDE), Montana, between 2004 and 2009 by following radio-collared females and observing their fate and reproductive performance. Our estimates of dependent cub and yearling survival were 0.612 (95% CI = 0.300–0.818) and 0.682 (95% CI = 0.258–0.898). Our estimates of subadult and
Authors
R.D. Mace, D.W. Carney, T. Chilton-Radandt, S.A. Courville, M.A. Haroldson, R.B. Harris, J. Jonkel, B. McLellan, M. Madel, T.L. Manley, C.C. Schwartz, C. Servheen, G. Stenhouse, J.S. Waller, E. Wenum