Mark Haroldson
Mark Haroldson is a USGS Supervisory Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT.
Research Interest
Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Education and Certifications
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Science and Products
Filter Total Items: 83
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2017
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2017. This report also contains a summary of grizzly bear management actions to address conflict situations. Annual reports of the IGBST summarize annual data collection. Because additional inform
Potential paths for male-mediated gene flow to and from an isolated grizzly bear population
For several decades, grizzly bear populations in the Greater Yellowstone Ecosystem (GYE) and the Northern Continental Divide Ecosystem (NCDE) have increased in numbers and range extent. The GYE population remains isolated and although effective population size has increased since the early 1980s, genetic connectivity between these populations remains a long-term management goal. With only ~110 km
Authors
Christopher P. Peck, Frank T. van Manen, Cecily M. Costello, Mark A. Haroldson, Lisa Landenburger, Lori L. Roberts, Daniel D. Bjornlie, Richard D. Mace
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2016
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2016. The report also contains a summary of grizzly bear management actions to address conflict situations.
Diet and macronutrient optimization in wild ursids: A comparison of grizzly bears with sympatric and allopatric black bears
When fed ad libitum, ursids can maximize mass gain by selecting mixed diets wherein protein provides 17 ± 4% of digestible energy, relative to carbohydrates or lipids. In the wild, this ability is likely constrained by seasonal food availability, limits of intake rate as body size increases, and competition. By visiting locations of 37 individuals during 274 bear-days, we documented foods consumed
Authors
Cecily M. Costello, Steven L. Cain, Shannon R Pils, Leslie Frattaroli, Mark A. Haroldson, Frank T. van Manen
Detecting grizzly bear use of ungulate carcasses using global positioning system telemetry and activity data
Global positioning system (GPS) wildlife collars have revolutionized wildlife research. Studies of predation by free-ranging carnivores have particularly benefited from the application of location clustering algorithms to determine when and where predation events occur. These studies have changed our understanding of large carnivore behavior, but the gains have concentrated on obligate carnivores.
Authors
Michael R. Ebinger, Mark A. Haroldson, Frank T. van Manen, Cecily M. Costello, Daniel D. Bjornlie, Daniel J. Thompson, Kerry A. Gunther, Jennifer K. Fortin, Justin E. Teisberg, Shannon R Pils, P J White, Steven L. Cain, Paul C. Cross
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2015
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2015. The report also contains a summary of grizzly bear management actions to address conflict situations.
Forty years of grizzly bear recovery in the Greater Yellowstone Ecosystem
No abstract available.
Authors
Frank T. van Manen, Cecily M. Costello, Kerry A. Gunther, Mark A. Haroldson
Density dependence, whitebark pine, and vital rates of grizzly bears
Understanding factors influencing changes in population trajectory is important for effective wildlife management, particularly for populations of conservation concern. Annual population growth of the grizzly bear (Ursus arctos) population in the Greater Yellowstone Ecosystem, USA has slowed from 4.2–7.6% during 1983–2001 to 0.3–2.2% during 2002–2011. Substantial changes in availability of a key f
Authors
Frank T. van Manen, Mark A. Haroldson, Daniel D. Bjornlie, Michael R. Ebinger, Daniel J. Thompson, Cecily M. Costello, Gary C. White
Multiple estimates of effective population size for monitoring a long-lived vertebrate: An application to Yellowstone grizzly bears
Effective population size (Ne) is a key parameter for monitoring the genetic health of threatened populations because it reflects a population's evolutionary potential and risk of extinction due to genetic stochasticity. However, its application to wildlife monitoring has been limited because it is difficult to measure in natural populations. The isolated and well-studied population of grizzly bea
Authors
Pauline L. Kamath, Mark A. Haroldson, Gordon Luikart, David Paetkau, Craig L. Whitman, Frank T. van Manen
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2014
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2014. The report also contains a summary of grizzly bear management actions to address conflict situations.
Whitebark pine, population density, and home-range size of grizzly bears in the greater Yellowstone ecosystem
Changes in life history traits of species can be an important indicator of potential factors influencing populations. For grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE), recent decline of whitebark pine (WBP; Pinus albicaulis), an important fall food resource, has been paired with a slowing of population growth following two decades of robust population increase. These obs
Authors
Daniel D. Bjornlie, Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Daniel J. Thompson, Cecily M. Costello
Science and Products
Filter Total Items: 83
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2017
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2017. This report also contains a summary of grizzly bear management actions to address conflict situations. Annual reports of the IGBST summarize annual data collection. Because additional inform
Potential paths for male-mediated gene flow to and from an isolated grizzly bear population
For several decades, grizzly bear populations in the Greater Yellowstone Ecosystem (GYE) and the Northern Continental Divide Ecosystem (NCDE) have increased in numbers and range extent. The GYE population remains isolated and although effective population size has increased since the early 1980s, genetic connectivity between these populations remains a long-term management goal. With only ~110 km
Authors
Christopher P. Peck, Frank T. van Manen, Cecily M. Costello, Mark A. Haroldson, Lisa Landenburger, Lori L. Roberts, Daniel D. Bjornlie, Richard D. Mace
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2016
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2016. The report also contains a summary of grizzly bear management actions to address conflict situations.
Diet and macronutrient optimization in wild ursids: A comparison of grizzly bears with sympatric and allopatric black bears
When fed ad libitum, ursids can maximize mass gain by selecting mixed diets wherein protein provides 17 ± 4% of digestible energy, relative to carbohydrates or lipids. In the wild, this ability is likely constrained by seasonal food availability, limits of intake rate as body size increases, and competition. By visiting locations of 37 individuals during 274 bear-days, we documented foods consumed
Authors
Cecily M. Costello, Steven L. Cain, Shannon R Pils, Leslie Frattaroli, Mark A. Haroldson, Frank T. van Manen
Detecting grizzly bear use of ungulate carcasses using global positioning system telemetry and activity data
Global positioning system (GPS) wildlife collars have revolutionized wildlife research. Studies of predation by free-ranging carnivores have particularly benefited from the application of location clustering algorithms to determine when and where predation events occur. These studies have changed our understanding of large carnivore behavior, but the gains have concentrated on obligate carnivores.
Authors
Michael R. Ebinger, Mark A. Haroldson, Frank T. van Manen, Cecily M. Costello, Daniel D. Bjornlie, Daniel J. Thompson, Kerry A. Gunther, Jennifer K. Fortin, Justin E. Teisberg, Shannon R Pils, P J White, Steven L. Cain, Paul C. Cross
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2015
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2015. The report also contains a summary of grizzly bear management actions to address conflict situations.
Forty years of grizzly bear recovery in the Greater Yellowstone Ecosystem
No abstract available.
Authors
Frank T. van Manen, Cecily M. Costello, Kerry A. Gunther, Mark A. Haroldson
Density dependence, whitebark pine, and vital rates of grizzly bears
Understanding factors influencing changes in population trajectory is important for effective wildlife management, particularly for populations of conservation concern. Annual population growth of the grizzly bear (Ursus arctos) population in the Greater Yellowstone Ecosystem, USA has slowed from 4.2–7.6% during 1983–2001 to 0.3–2.2% during 2002–2011. Substantial changes in availability of a key f
Authors
Frank T. van Manen, Mark A. Haroldson, Daniel D. Bjornlie, Michael R. Ebinger, Daniel J. Thompson, Cecily M. Costello, Gary C. White
Multiple estimates of effective population size for monitoring a long-lived vertebrate: An application to Yellowstone grizzly bears
Effective population size (Ne) is a key parameter for monitoring the genetic health of threatened populations because it reflects a population's evolutionary potential and risk of extinction due to genetic stochasticity. However, its application to wildlife monitoring has been limited because it is difficult to measure in natural populations. The isolated and well-studied population of grizzly bea
Authors
Pauline L. Kamath, Mark A. Haroldson, Gordon Luikart, David Paetkau, Craig L. Whitman, Frank T. van Manen
Yellowstone grizzly bear investigations — Annual report of the Interagency Grizzly Bear Study Team 2014
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2014. The report also contains a summary of grizzly bear management actions to address conflict situations.
Whitebark pine, population density, and home-range size of grizzly bears in the greater Yellowstone ecosystem
Changes in life history traits of species can be an important indicator of potential factors influencing populations. For grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE), recent decline of whitebark pine (WBP; Pinus albicaulis), an important fall food resource, has been paired with a slowing of population growth following two decades of robust population increase. These obs
Authors
Daniel D. Bjornlie, Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Daniel J. Thompson, Cecily M. Costello