Mark Haroldson
Mark Haroldson is a USGS Supervisory Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT.
Research Interest
Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Education and Certifications
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Science and Products
Filter Total Items: 83
Use of isotopic sulfur to determine whitebark pine consumption by Yellowstone bears: a reassessment
Use of naturally occurring stable isotopes to estimate assimilated diet of bears is one of the single greatest breakthroughs in nutritional ecology during the past 20 years. Previous research in the Greater Yellowstone Ecosystem (GYE), USA, established a positive relationship between the stable isotope of sulfur (δ34S) and consumption of whitebark pine (Pinus albicaulis) seeds. That work combined
Authors
Charles C. Schwartz, Justin E. Teisberg, Jennifer K. Fortin, Mark A. Haroldson, Christopher Servheen, Charles T. Robbins, Frank T. van Manen
Roadside bear viewing opportunities in Yellowstone National Park: characteristics, trends, and influence of whitebark pine
Opportunities for viewing grizzly bears (Ursus arctos) and American black bears (U. americanus) from roadways in Yellowstone National Park (YNP) have increased in recent years. Unlike the panhandling bears common prior to the 1970s, current viewing usually involves bears feeding on natural foods. We define roadside bear viewing opportunities that cause traffic congestion as ‘‘bear-jams.’’ We inves
Authors
Mark A. Haroldson, Kerry Gunther
Re-evaluation of Yellowstone grizzly bear population dynamics not supported by empirical data: response to Doak & Cutler
Doak and Cutler critiqued methods used by the Interagency Grizzly Bear Study Team (IGBST) to estimate grizzly bear population size and trend in the Greater Yellowstone Ecosystem. Here, we focus on the premise, implementation, and interpretation of simulations they used to support their arguments. They argued that population increases documented by IGBST based on females with cubs-of-the-year were
Authors
Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Richard B. Harris, Megan D. Higgs, Steve Cherry, Gary C. White, Charles C. Schwartz
Influence of whitebark pine decline on fall habitat use and movements of grizzly bears in the Greater Yellowstone Ecosystem
When abundant, seeds of the high-elevation whitebark pine (WBP; Pinus albicaulis) are an important fall food for grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem. Rates of bear mortality and bear/human conflicts have been inversely associated with WBP productivity. Recently, mountain pine beetles (Dendroctonus ponderosae) have killed many cone-producing WBP trees. We used fall (15
Authors
Cecily M. Costello, Frank T. van Manen, Mark A. Haroldson, Michael R. Ebinger, Steven L. Cain, Kerry A. Gunther, Daniel D. Bjornlie
Methods to estimate distribution and range extent of grizzly bears in the Greater Yellowstone Ecosystem
The distribution of the Greater Yellowstone Ecosystem grizzly bear (Ursus arctos) population has expanded into areas unoccupied since the early 20th century. Up-to-date information on the area and extent of this distribution is crucial for federal, state, and tribal wildlife and land managers to make informed decisions regarding grizzly bear management. The most recent estimate of grizzly bear dis
Authors
Mark A. Haroldson, Charles C. Schwartz, Daniel J. Thompson, Daniel D. Bjornlie, Kerry A. Gunther, Steven L. Cain, Daniel B. Tyers, Kevin L. Frey, Bryan C. Aber
Contrasting past and current numbers of bears visiting Yellowstone cutthroat trout streams
Spawning cutthroat trout (Oncorhynchus clarkii bouvieri) were historically abundant within tributary streams of Yellowstone Lake within Yellowstone National Park and were a highly digestible source of energy and protein for Yellowstone’s grizzly bears (Ursus arctos) and black bears (U. americanus). The cutthroat trout population has subsequently declined since the introduction of non-native lake t
Authors
Mark A. Haroldson, Charles C. Schwartz, Justin E. Teisberg, Kerry A. Gunther, Jennifer K. Fortin, Charles T. Robbins
Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2013. The report also contains a summary of nuisance grizzly bear management actions.
Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem
Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE) are opportunistic omnivores that eat a great diversity of plant and animal species. Changes in climate may affect regional vegetation, hydrology, insects, and fire regimes, likely influencing the abundance, range, and elevational distribution of the plants and animals consumed by GYE grizzly bears. Determining the dietary brea
Authors
Kerry A. Gunther, Rebecca Shoemaker, Kevin L. Frey, Mark A. Haroldson, Steven L. Cain, Frank T. van Manen, Jennifer K. Fortin
Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee
The Yellowstone grizzly bear (Ursus arctos) was listed as a threatened species in 1975 (Federal Register 40 FR:31734-31736). Since listing, recovery efforts have focused on increasing population size, improving habitat security, managing bear mortalities, and reducing bear-human conflicts. The Interagency Grizzly Bear Committee (IGBC; partnership of federal and state agencies responsible for grizz
Authors
Frank T. van Manen, Cecily M. Costello, Mark A. Haroldson, Daniel D. Bjornlie, Michael R. Ebinger, Kerry A. Gunther, Mary Frances Mahalovich, Daniel J. Thompson, Megan D. Higgs, Kathryn M. Irvine, Kristin Legg, Daniel B. Tyers, Lisa Landenburger, Steven L. Cain, Kevin L. Frey, Bryan C. Aber, Charles C. Schwartz
Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year
Mark-resight designs for estimation of population abundance are common and attractive to researchers. However, inference from such designs is very limited when faced with sparse data, either from a low number of marked animals, a low probability of detection, or both. In the Greater Yellowstone Ecosystem, yearly mark-resight data are collected for female grizzly bears with cubs-of-the-year (FCOY),
Authors
Megan D. Higgs, William A. Link, Gary C. White, Mark A. Haroldson, Daniel D. Bjornlie
Omnivory and the terrestrial food web: Yellowstone grizzly beard diets
No abstract available.
Authors
Charles C. Schwartz, Mark A. Haroldson, Kerry A. Gunther, Charles T. Robbins
Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem
The Greater Yellowstone Ecosystem (GYE) has experienced changes in the distribution and availability of grizzly bear (Ursus arctos) food resources in recent decades. The decline of ungulates, fish, and whitebark pine seeds (Pinus albicaulis) has prompted questions regarding their ability to adapt. We examined body composition and diet of grizzly bears using bioelectrical impedance and stable isoto
Authors
Charles C. Schwartz, Jennifer K. Fortin, Justin E. Teisberg, Mark A. Haroldson, Christopher Servheen, Charles T. Robbins, Frank T. van Manen
Science and Products
Filter Total Items: 83
Use of isotopic sulfur to determine whitebark pine consumption by Yellowstone bears: a reassessment
Use of naturally occurring stable isotopes to estimate assimilated diet of bears is one of the single greatest breakthroughs in nutritional ecology during the past 20 years. Previous research in the Greater Yellowstone Ecosystem (GYE), USA, established a positive relationship between the stable isotope of sulfur (δ34S) and consumption of whitebark pine (Pinus albicaulis) seeds. That work combined
Authors
Charles C. Schwartz, Justin E. Teisberg, Jennifer K. Fortin, Mark A. Haroldson, Christopher Servheen, Charles T. Robbins, Frank T. van Manen
Roadside bear viewing opportunities in Yellowstone National Park: characteristics, trends, and influence of whitebark pine
Opportunities for viewing grizzly bears (Ursus arctos) and American black bears (U. americanus) from roadways in Yellowstone National Park (YNP) have increased in recent years. Unlike the panhandling bears common prior to the 1970s, current viewing usually involves bears feeding on natural foods. We define roadside bear viewing opportunities that cause traffic congestion as ‘‘bear-jams.’’ We inves
Authors
Mark A. Haroldson, Kerry Gunther
Re-evaluation of Yellowstone grizzly bear population dynamics not supported by empirical data: response to Doak & Cutler
Doak and Cutler critiqued methods used by the Interagency Grizzly Bear Study Team (IGBST) to estimate grizzly bear population size and trend in the Greater Yellowstone Ecosystem. Here, we focus on the premise, implementation, and interpretation of simulations they used to support their arguments. They argued that population increases documented by IGBST based on females with cubs-of-the-year were
Authors
Frank T. van Manen, Michael R. Ebinger, Mark A. Haroldson, Richard B. Harris, Megan D. Higgs, Steve Cherry, Gary C. White, Charles C. Schwartz
Influence of whitebark pine decline on fall habitat use and movements of grizzly bears in the Greater Yellowstone Ecosystem
When abundant, seeds of the high-elevation whitebark pine (WBP; Pinus albicaulis) are an important fall food for grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem. Rates of bear mortality and bear/human conflicts have been inversely associated with WBP productivity. Recently, mountain pine beetles (Dendroctonus ponderosae) have killed many cone-producing WBP trees. We used fall (15
Authors
Cecily M. Costello, Frank T. van Manen, Mark A. Haroldson, Michael R. Ebinger, Steven L. Cain, Kerry A. Gunther, Daniel D. Bjornlie
Methods to estimate distribution and range extent of grizzly bears in the Greater Yellowstone Ecosystem
The distribution of the Greater Yellowstone Ecosystem grizzly bear (Ursus arctos) population has expanded into areas unoccupied since the early 20th century. Up-to-date information on the area and extent of this distribution is crucial for federal, state, and tribal wildlife and land managers to make informed decisions regarding grizzly bear management. The most recent estimate of grizzly bear dis
Authors
Mark A. Haroldson, Charles C. Schwartz, Daniel J. Thompson, Daniel D. Bjornlie, Kerry A. Gunther, Steven L. Cain, Daniel B. Tyers, Kevin L. Frey, Bryan C. Aber
Contrasting past and current numbers of bears visiting Yellowstone cutthroat trout streams
Spawning cutthroat trout (Oncorhynchus clarkii bouvieri) were historically abundant within tributary streams of Yellowstone Lake within Yellowstone National Park and were a highly digestible source of energy and protein for Yellowstone’s grizzly bears (Ursus arctos) and black bears (U. americanus). The cutthroat trout population has subsequently declined since the introduction of non-native lake t
Authors
Mark A. Haroldson, Charles C. Schwartz, Justin E. Teisberg, Kerry A. Gunther, Jennifer K. Fortin, Charles T. Robbins
Yellowstone grizzly bear investigations—Annual report of the Interagency Grizzly Bear Study Team 2013
This Annual Report summarizes results of grizzly bear (Ursus arctos) monitoring and research conducted in the Greater Yellowstone Ecosystem (GYE) by the Interagency Grizzly Bear Study Team (IGBST) during 2013. The report also contains a summary of nuisance grizzly bear management actions.
Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem
Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE) are opportunistic omnivores that eat a great diversity of plant and animal species. Changes in climate may affect regional vegetation, hydrology, insects, and fire regimes, likely influencing the abundance, range, and elevational distribution of the plants and animals consumed by GYE grizzly bears. Determining the dietary brea
Authors
Kerry A. Gunther, Rebecca Shoemaker, Kevin L. Frey, Mark A. Haroldson, Steven L. Cain, Frank T. van Manen, Jennifer K. Fortin
Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee
The Yellowstone grizzly bear (Ursus arctos) was listed as a threatened species in 1975 (Federal Register 40 FR:31734-31736). Since listing, recovery efforts have focused on increasing population size, improving habitat security, managing bear mortalities, and reducing bear-human conflicts. The Interagency Grizzly Bear Committee (IGBC; partnership of federal and state agencies responsible for grizz
Authors
Frank T. van Manen, Cecily M. Costello, Mark A. Haroldson, Daniel D. Bjornlie, Michael R. Ebinger, Kerry A. Gunther, Mary Frances Mahalovich, Daniel J. Thompson, Megan D. Higgs, Kathryn M. Irvine, Kristin Legg, Daniel B. Tyers, Lisa Landenburger, Steven L. Cain, Kevin L. Frey, Bryan C. Aber, Charles C. Schwartz
Insights into the latent multinomial model through mark-resight data on female grizzly bears with cubs-of-the-year
Mark-resight designs for estimation of population abundance are common and attractive to researchers. However, inference from such designs is very limited when faced with sparse data, either from a low number of marked animals, a low probability of detection, or both. In the Greater Yellowstone Ecosystem, yearly mark-resight data are collected for female grizzly bears with cubs-of-the-year (FCOY),
Authors
Megan D. Higgs, William A. Link, Gary C. White, Mark A. Haroldson, Daniel D. Bjornlie
Omnivory and the terrestrial food web: Yellowstone grizzly beard diets
No abstract available.
Authors
Charles C. Schwartz, Mark A. Haroldson, Kerry A. Gunther, Charles T. Robbins
Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem
The Greater Yellowstone Ecosystem (GYE) has experienced changes in the distribution and availability of grizzly bear (Ursus arctos) food resources in recent decades. The decline of ungulates, fish, and whitebark pine seeds (Pinus albicaulis) has prompted questions regarding their ability to adapt. We examined body composition and diet of grizzly bears using bioelectrical impedance and stable isoto
Authors
Charles C. Schwartz, Jennifer K. Fortin, Justin E. Teisberg, Mark A. Haroldson, Christopher Servheen, Charles T. Robbins, Frank T. van Manen