Mark Haroldson
Mark Haroldson is a USGS Supervisory Wildlife Biologist for the Interagency Grizzly Bears Study Team and works out of the Northern Rocky Mountain Science Center, Bozeman, MT.
Research Interest
Mark has had a passion for bears since he began studying them as an undergraduate in 1976. He has worked in bear research and management in several western states ever since. Since 1984 he has worked in various capacities for the Interagency Grizzly Bear Study Team in the Greater Yellowstone Ecosystem. Current research is focused on mortality and population trend of grizzly bears in the Greater Yellowstone Ecosystem.
Education and Certifications
B.S. Wildlife Biology, 1979, University of Montana
Graduate level class work, 1982-88, University of Montana
Science and Products
Filter Total Items: 83
Large carnivores response to recreational big game hunting along the Yellowstone National Park and Absaroka-Beartooth Wilderness boundary
The Greater Yellowstone Ecosystem contains the rare combination of an intact guild of native large carnivores, their prey, and differing land management policies (National Park versus National Forest; no hunting versus hunting). Concurrent field studies on large carnivores allowed us to investigate activities of humans and carnivores on Yellowstone National Park's (YNP) northern boundary. Prior to
Authors
T.E. Ruth, D.W. Smith, M.A. Haroldson, P.C. Buotte, C.C. Schwartz, H.B. Quigley, S. Cherry, D. Tyres, K. Frey
Grizzly bear
The grizzly bear inspires fear, awe, and respect in humans to a degree unmatched by any other North American wild mammal. Like other bear species, it can inflict serious injury and death on humans and sometimes does. Unlike the polar bear (Ursus maritimus) of the sparsely inhabited northern arctic, however, grizzly bears still live in areas visited by crowds of people, where presence of the grizzl
Authors
C.C. Schwartz, S.D. Miller, M.A. Haroldson
Yellowstone grizzly bear investigations: Annual report of the Interagency Grizzly Bear Study Team, 2001
The contents of this Annual Report summarize results of monitoring and research from the 2001 field season. The report also contains a summary of nuisance grizzly bear (Ursus arctos horribilis) management actions.
In addition to our normal monitoring, we completed an array of studies addressing the potential impacts of winter recreation on denning grizzly bears. This research was in response to a
Estimating total human-caused mortality from reported mortality using data from radio-instrumented grizzly bears
Tracking mortality of the Yellowstone grizzly bear (Ursus arctos horribilis) is an essential issue of the recovery process. Problem bears removed by agencies are well documented. Deaths of radiocollared bears are known or, in many cases, can be reliably inferred. Additionally, the public reports an unknown proportion of deaths of uncollared bears. Estimating the number of non-agency human-caused
Authors
S. Cherry, M.A. Haroldson, J. Robison-Cox, C.C. Schwartz
Grizzly bear denning chronology and movements in the Greater Yellowstone Ecosystem
Den entrance and emergence dates of grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem are important to management agencies that wish to minimize impacts of human activities on bears. Current estimates for grizzly bear denning events use data that were collected from 1975–80. We update these estimates by including data obtained from 1981–99. We used aerial telemetry data to estimate
Authors
Mark A. Haroldson, Mark A. Ternent, Kerry A. Gunther, Charles C. Schwartz
Consumption of fungal sporocarps by Yellowstone grizzly bears
Sign of grizzly bears (Ursus arctos horribilis) consuming fungal sporocarps (mushrooms and truffles) was observed on 68 occasions during a study of radiomarked bears in the Yellowstone region, 1977–96. Sporocarps also were detected in 96 grizzly bear feces. Most fungi consumedby Yellowstone's grizzlybearsweremembersofthe Boletaceae(Suillus spp.), Russulaceae (Russula spp. and Lactarius sp.), Morc
Authors
David J. Mattson, Shannon Podruzny, Mark A. Haroldson
Estimating the number of females with cubs-of-the-year in the Yellowstone grizzly bear population
No abstract available.
Authors
K.A. Keating, C.C. Schwartz, M.A. Haroldson, D. Moody
Distribution of grizzly bears in the Greater Yellowstone Ecosystem, 1990-2000
The Yellowstone grizzly bear (Ursus arctos horribilis) has been expanding its range during the past 2 decades and now occupies historic habitats that had been vacant. A current understanding of the distribution of grizzly bears within the ecosystem is useful in the recovery process and to help guide the state and federal land management agencies and state wildlife agencies of Idaho, Wyoming, and M
Authors
Charles C. Schwartz, Mark A. Haroldson, Kerry A. Gunther, D. Moody
Negative binomial models for abundance estimation of multiple closed populations
Counts of uniquely identified individuals in a population offer opportunities to estimate abundance. However, for various reasons such counts may be burdened by heterogeneity in the probability of being detected. Theoretical arguments and empirical evidence demonstrate that the negative binomial distribution (NBD) is a useful characterization for counts from biological populations with heterogenei
Authors
Mark S. Boyce, Darry I. MacKenzie, Bryan F.J. Manly, Mark A. Haroldson, David W. Moody
Effects of exotic species on Yellowstone's grizzly bears
Humans have affected grizzly bears (Ursus arctos horribilis) by direct mortality, competition for space and resources, and introduction of exotic species. Exotic organisms that have affected grizzly bears in the Greater Yellowstone Area include common dandelion (Taraxacum officinale), nonnative clovers (Trifolium spp.), domesticated livestock, bovine brucellosis (Brucella abortus), lake trout (Sal
Authors
Daniel P. Reinhart, Mark A. Haroldson, D.J. Mattson, Kerry A. Gunther
Estimating numbers of females with cubs-of-the-year in the Yellowstone grizzly bear population
For grizzly bears (Ursus arctos horribilis) in the Greater Yellowstone Ecosystem (GYE), minimum population size and allowable numbers of human-caused mortalities have been calculated as a function of the number of unique females with cubs-of-the-year (FCUB) seen during a 3- year period. This approach underestimates the total number of FCUB, thereby biasing estimates of population size and sustaina
Authors
K.A. Keating, C.C. Schwartz, M.A. Haroldson, D. Moody
Science and Products
Filter Total Items: 83
Large carnivores response to recreational big game hunting along the Yellowstone National Park and Absaroka-Beartooth Wilderness boundary
The Greater Yellowstone Ecosystem contains the rare combination of an intact guild of native large carnivores, their prey, and differing land management policies (National Park versus National Forest; no hunting versus hunting). Concurrent field studies on large carnivores allowed us to investigate activities of humans and carnivores on Yellowstone National Park's (YNP) northern boundary. Prior to
Authors
T.E. Ruth, D.W. Smith, M.A. Haroldson, P.C. Buotte, C.C. Schwartz, H.B. Quigley, S. Cherry, D. Tyres, K. Frey
Grizzly bear
The grizzly bear inspires fear, awe, and respect in humans to a degree unmatched by any other North American wild mammal. Like other bear species, it can inflict serious injury and death on humans and sometimes does. Unlike the polar bear (Ursus maritimus) of the sparsely inhabited northern arctic, however, grizzly bears still live in areas visited by crowds of people, where presence of the grizzl
Authors
C.C. Schwartz, S.D. Miller, M.A. Haroldson
Yellowstone grizzly bear investigations: Annual report of the Interagency Grizzly Bear Study Team, 2001
The contents of this Annual Report summarize results of monitoring and research from the 2001 field season. The report also contains a summary of nuisance grizzly bear (Ursus arctos horribilis) management actions.
In addition to our normal monitoring, we completed an array of studies addressing the potential impacts of winter recreation on denning grizzly bears. This research was in response to a
Estimating total human-caused mortality from reported mortality using data from radio-instrumented grizzly bears
Tracking mortality of the Yellowstone grizzly bear (Ursus arctos horribilis) is an essential issue of the recovery process. Problem bears removed by agencies are well documented. Deaths of radiocollared bears are known or, in many cases, can be reliably inferred. Additionally, the public reports an unknown proportion of deaths of uncollared bears. Estimating the number of non-agency human-caused
Authors
S. Cherry, M.A. Haroldson, J. Robison-Cox, C.C. Schwartz
Grizzly bear denning chronology and movements in the Greater Yellowstone Ecosystem
Den entrance and emergence dates of grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem are important to management agencies that wish to minimize impacts of human activities on bears. Current estimates for grizzly bear denning events use data that were collected from 1975–80. We update these estimates by including data obtained from 1981–99. We used aerial telemetry data to estimate
Authors
Mark A. Haroldson, Mark A. Ternent, Kerry A. Gunther, Charles C. Schwartz
Consumption of fungal sporocarps by Yellowstone grizzly bears
Sign of grizzly bears (Ursus arctos horribilis) consuming fungal sporocarps (mushrooms and truffles) was observed on 68 occasions during a study of radiomarked bears in the Yellowstone region, 1977–96. Sporocarps also were detected in 96 grizzly bear feces. Most fungi consumedby Yellowstone's grizzlybearsweremembersofthe Boletaceae(Suillus spp.), Russulaceae (Russula spp. and Lactarius sp.), Morc
Authors
David J. Mattson, Shannon Podruzny, Mark A. Haroldson
Estimating the number of females with cubs-of-the-year in the Yellowstone grizzly bear population
No abstract available.
Authors
K.A. Keating, C.C. Schwartz, M.A. Haroldson, D. Moody
Distribution of grizzly bears in the Greater Yellowstone Ecosystem, 1990-2000
The Yellowstone grizzly bear (Ursus arctos horribilis) has been expanding its range during the past 2 decades and now occupies historic habitats that had been vacant. A current understanding of the distribution of grizzly bears within the ecosystem is useful in the recovery process and to help guide the state and federal land management agencies and state wildlife agencies of Idaho, Wyoming, and M
Authors
Charles C. Schwartz, Mark A. Haroldson, Kerry A. Gunther, D. Moody
Negative binomial models for abundance estimation of multiple closed populations
Counts of uniquely identified individuals in a population offer opportunities to estimate abundance. However, for various reasons such counts may be burdened by heterogeneity in the probability of being detected. Theoretical arguments and empirical evidence demonstrate that the negative binomial distribution (NBD) is a useful characterization for counts from biological populations with heterogenei
Authors
Mark S. Boyce, Darry I. MacKenzie, Bryan F.J. Manly, Mark A. Haroldson, David W. Moody
Effects of exotic species on Yellowstone's grizzly bears
Humans have affected grizzly bears (Ursus arctos horribilis) by direct mortality, competition for space and resources, and introduction of exotic species. Exotic organisms that have affected grizzly bears in the Greater Yellowstone Area include common dandelion (Taraxacum officinale), nonnative clovers (Trifolium spp.), domesticated livestock, bovine brucellosis (Brucella abortus), lake trout (Sal
Authors
Daniel P. Reinhart, Mark A. Haroldson, D.J. Mattson, Kerry A. Gunther
Estimating numbers of females with cubs-of-the-year in the Yellowstone grizzly bear population
For grizzly bears (Ursus arctos horribilis) in the Greater Yellowstone Ecosystem (GYE), minimum population size and allowable numbers of human-caused mortalities have been calculated as a function of the number of unique females with cubs-of-the-year (FCUB) seen during a 3- year period. This approach underestimates the total number of FCUB, thereby biasing estimates of population size and sustaina
Authors
K.A. Keating, C.C. Schwartz, M.A. Haroldson, D. Moody